In the aftermath of the final events of Velosaura, Qora races to the independent peninsula known as the Tail, where enormous dinosaurs exist in harmony with people who live high in the trees. With her brother's help, she must determine whether it will be possible to harness these dinosaurs for battle-and persuade the people of the Tail to help her do it.

After a turbulent wedding ceremony, Ninan and Paqari are close behind, ready to join forces with the Allpan quya, the Razorclaws, and a few new allies to prepare for the oncoming war.

When Qhapaq Apo reveals new reptiles whose abilities verge on the fantastic, all branches of the resistance must come together to defeat these otherworldly monsters once and for all.